Prof. Dr. Rolf Beutel

 

 

Friedrich-Schiller-Universität Jena
Institut für Zoologie und Evolutionsforschung
mit Phyletischem Museum, Ernst-Haeckel-Haus und Biologiedidaktik
Erbertstr. 1
D-07743 Jena

Fon:

03641/ 94 91 53

Fax:

03641/ 94 91 42

 

Studium in Tübingen, Dissertation bei Dr. G. Mickoleit am Institut für Biologie III, Titel: Skelet und Muskulatur des Kopfes und Thorax von Hygrobia tarda (Herbst). Ein Beitrag zur Klärung der phylogene­tischen Beziehungen der Hydradephaga (Insecta: Coleoptera). Abschluss 1986

1986-1987: Postdoctoral fellowship, University of Manitoba, Dept. of Entomo­logy, Winnipeg, Canada

1987-1994: Assistent am Insti­tut für Biologie II (Zoologie), RWTH Aachen

1994-1995: Assistent am Insti­tut für spezielle Zoologie und Evolutionsbiologie, FSU Jena

Habilitation: Dezember 1995

1996-2004: Privatdozent am Insti­tut für spezielle Zoologie und Evolutionsbiologie, FSU Jena

1999: Ruf auf eine Professur für Entomologie an der Kansas State University

2004: Ruf auf eine C4-Professur für Zoologie (Entomologie) an der Universität Hamburg

Seit 2004 Professor am Institut für Spezielle Zoologie und Evolutionsbiologie und verantwortlich für die wissenschaftliche Sammlung am Museum


Topic01: The Evolution of Endopterygota - reconstructing the phylogeny of the most successful lineage of organisms with phylogenomic and morphological data (DFG BE1789/8-1)

Topic02: 1KITE, Reconstructing the phylogeny of Hexapoda using transcriptomes

Topic03: The phylogeny and evolution of the "Lower Neoptera" (Insecta) (DFG BE1789/10-1)

Topic04: Der männliche Genitalapparat der holometabolen Insekten und seine evolutive Bedeutung (VolkswagenStiftung)

Topic05: The evolution of larval and adult features in Diptera (Insecta) (DFG 1789/6-1)

Topic06:  The phylogeny of the megadiverse Chrysomeloidea using morpholocical characters and molecular data (Chinese Academy of Sciences, Beijing)

Topic07: The evolution of attachment structures in Paraneoptera (IMPRS Jena)

Topic08: Miniaturisation in insects

Topic09: Innovative morphological techniques

Topic10: Extension and modification of Morph D Base producing a system for permanent storage and documentation of volume data of biological objects in high resolution (DFG BE1789/9-1)


 

Topic01: The Evolution of Endopterygota

Co-operation with Prof. Dr. B. Misof (Zoologisches Forschungsmuseum Alexander König, Bonn)

Endopterygota (=Holometabola) are the most successful lineage of living organisms (ca. 800.000). Even though strong efforts have been made recently to reconstruct the phylogenetic relationships, different molecular data sets and morphological characters still yield conflicting results. The evolution of the group is far from being understood. For the reconstruction of a robust backbone of the holometabolan phylogeny extensive sequence data will be compiled (ESTs and single gene data) and analysed separately and in combination with large morphological data sets from an earlier project (BE 1789/4-1; e.g., Beutel et al. 2010). The branching pattern will enable us to reconstruct the evolution of different character complexes, such as for instance larval head structures associated with feeding, or the thoracic locomotor apparatus. Scenarios for the early evolution of holometabolous insects will be developed.

The main issues are 1) The interrelationships of the holometabolan orders; 2) The evolution of different character complexes and the groundplan of the lineage; 3) The assessment of data quality and information content of molecular and morphological data.

 

Topic02: 1KITE, Reconstructing the phylogeny of Hexapoda using transcriptomes

Co-operation with Prof. Dr. B. Misof (Zoologisches Forschungsmuseum Alexander König, Bonn), Prof. Dr. K. Kjer (Rutgers University, NJ), Dr. X. Zhou (Beijing Genomics Institute, Shenzhen), Prof. Dr. R. Machida (University of Tsukuba, Sugadaira Mountain Research Centre), Prof. Dr. G. Pass (Universität Wien), Dr. A. Stamatakis (Heidelberg Institute for Theoretical Studies), Dr. Frank Friedrich (Universität Hamburg) and others

Insects are by far the most species rich metazoan group and include many model organisms and economically and medically relevant species. Nevertheless, compared to vertebrates, insect transcriptomes have not been explored in depth for scientific and applied questions. The 1KITE initiative will alleviate this problem by characterizing 1000 transcriptomes of selected insect species covering the full taxonomic range, from ancestrally wingless hexapods to Holometabola. Special focus will be on transcriptomes of key taxa, including environmentally, economically and medically important groups. Several subprojects (e.g., Polyneoptera, Paraneoptera, Antliophora, Hymenoptera) will be carried in the framework of the project. The main goal aim is to deliver a robust backbone tree for the entire Hexapoda. Additionally, fully characterized transcriptomes of insects with impact on humans will be made available.

 

Topic03: The phylogeny and evolution of the "Lower Neoptera" (Insecta)

Dipl. Biol. Benjamin Wipfler, co-operation with PD. Dr. T. Hörnschemeyer, Dr. S. Bradler (Georg-August-Universität Göttingen), Dr.  K. Klass (Museum für Tierkunde, Senckenberg Natural History Collections Dresden)

Even though great efforts have been made to reconstruct the phylogeny of the lower neopteran insects (Polyneoptera), their relationships are still largely unresolved. This is presently the greatest gap in the understanding of insect systematics and evolution. The "Lower Neoptera" comprise nine lineages, the species-rich Orthoptera and Dictyoptera, the Plecoptera, the highly specialised Phasmatodea, Embioptera, and Dermaptera, and the very small orders Grylloblattodea, Mantophasmatodea, and Zoraptera. The knowledge of the morphology is fragmentary and superficial. Our project is designed to compile an extensive morphological dataset with a focus on presumptive key taxa. The phylogenetic reconstruction will provide a source for testing molecular trees and for placing extinct taxa (see e.g., 1KITE). A well supported phylogenetic hypothesis and comprehensive evolutionary scenario for the "Lower Neoptera" are the ultimate goals of the project.

 

Topic04: Der männliche Genitalapparat der holometabolen Insekten und seine evolutive Bedeutung/The male genital apparatus of holometabolous insects and its evolutionary significance

Dr. Frank Hünefeld, co-operation with Prof. Dr. R. Dallai (University of Siena), Prof. Dr. R. Machida (University of Tsukuba) and Prof. Dr. R. Meier (National University of Singapore)

The project combines aspects of museum specific research, evolution, phylogeny, and taxonomy. Its focus is on the male genital apparatus. With its complexity and extreme variability it has likely played a very important role in the evolution of the megadiverse insects. The 'lock-and-key mechanism' was often addressed by evolutionary biologists in the context of speciation processes. We apply innovative techniques to investigate external and internal genital structures. The largely unclear homology of elements of the male apparatus is addressed in a phylogenetic context. Genital characters will be coded and analysed with other morphological and molecular data (see Endopterygota project and 1KITE). Based on the results an evolutionary scenario for the male genital apparatus of Holometabola will be developed. An exemplar group (genus Eois, Geometridae, Lepidoptera) was chosen to optimise the study of genitalia in the context of taxonomic revisions with museum material. The significance of the 'lock-and-key mechanism' and of sexual selection will be critically evaluated.  A workshop will be carried out with a focus on innovative morphological techniques (2012). Models based on computer-reconstructions and results of the project will be presented in an exhibition ("Insects and Sex") in the Phyletisches Museum.

 

Topic05: The evolution of larval and adult features in Diptera (Insecta)

Dipl. Biol. Katharina Schneeberg, Dipl. Biol. Benjamin Wipfler, co-operation with Prof. Dr. G. Courtney (Iowa State University) and others

Diptera are an extremely species rich group with ca. 150.000 species. As vectors of numerous diseases adults have unparalleled negative impacts on human health (e.g., Malaria [Anopheles]). Head structures play a major role in this context. The knowledge of adult and larval anatomy in general is very fragmentary. Therefore the first step will be a documentation of morphological data using innovative techniques (including amber fossils) (1). An extensive morphological character set will be compiled (adults and larvae) (2). The phylogenetic evaluation will also include molecular data (Flytree project [NSF]) (3). An evolutionary scenario will be developed, with emphasis on the adult head and feeding apparatus and larval features and habits (4). Groundplans (Diptera and Brachycera) will be reconstructed. The position of Diptera will be re-assessed using data from 'Phylogeny and Evolution of Endopterygota' (BE 1789/4-1). A subproject is a documentation of the morphology of Drosophila with innovative techniques. Shared expertise, data and material will guarantee maximum efficiency.    

 

Topic06:  The phylogeny of the megadiverse Chrysomeloidea using morpholocical characters and molecular data

Co-operation with Assoc. Prof. Dr. S.-Q. Ge, Prof. Dr. X.-K. Yang (Chinese Academy of Sciences, Beijing) and Prof. Dr. A. Ślipiński (CSIRO Canberra)

Chrysomeloidea are a megadiverse lineage with ca. 50.000 described species. Many members of the group are economically important as pests of cultivated plants. The phylogenetic relationships on the subfamily level will be evaluated with a carefully chosen taxon sampling and a broad spectrum of characters of all life stages (e.g., skeleto-muscular system, wing articulation, attachment structures). Extensive material of larvae, pupae and adults is already available at the Institute of Zoology of the Chinese Academy of Sciences (Beijing) and additional critical taxa will be provided by the Australian Insect Collection (ANIC, CSIRO Canberra).

 

Topic07: The evolution of attachment structures in Paraneoptera (International Max Planck Research School, MPI for Chemical Ecology)

Dipl. Biol. Katrin Friedemann, Co-operation with Max Planck Institute for Chemical Ecology (Jena) and Prof. Dr. S. Gorb (Universität Kiel)

Functional and evolutionary aspects of attachment structures in Paraneoptera (lice, cicada, true bugs etc.) are investigated.

 

Topic08: Miniaturisation

Co-operation with Dr. hab. H. Pohl (FSU Jena) and Dr. Alexey Polilov (Moscov State University)

The effects of miniaturisation in different groups (e.g., Strepsiptera, Hmenoptera, Diptera, Ptiliidae [Coleoptera]) and on different organ systems (e.g., musculature, cerebrum) are studied. Applied techniques comprise SEM, semithin histological sections, TEM, µ-CT, and 3-dimensional computer reconstruction. 

 

Topic09: Innovative morphological techniques

Co-operation with Dr. Felix Beckmann (DESY Hamburg) and others

A central aspect of our work is to develop and optimize innovative morphological techniques. In our phylogenetic projects we use different combinations of histology, SEM, 3D reconstruction, CLSM and x-ray micro-computertomography. An optimized workflow allows a greatly accelerated acquisition of high quality anatomical data with a minimum of material.

 

Topic10: Extension and modification of MorphDBase producing a system for permanent storage and documentation of volume data of biological objects in high resolution (DFG BE1789/9-1) (LIS Förderprogramm "Informationsmanagement")

Co-operation with Dr. L. Vogt, Prof. Dr. J. Wägele (Zoologisches Forschungsmuseum Alexander König Bonn), PD Dr. T. Hörnschemeyer (Georg-August-Universität Göttingen) and Prof. Dr. H. Hoch (Naturkundemuseum Berlin)

The aim of this project is to create optimal storage facilities for morphological megadata, based on an extended version of MorphDBase. Morphological results obtained with different techniques will be made accessible to the scientific community to maximise synergy effects and enhance research in evolutionary morphology and phylogeny.

 


 




Mitgliedschaft in Gremien:

Fakultätsrat der Fakultät für Biologie und Pharmazie der FSU Jena (seit 2005)
Haushaltsausschuss des Fakultätsrats (seit 2005)
Tierschutzkomittee des Landes Thüringen
Stiftungsrat der Stiftung Naturschutz Thüringen

Editorial advisor:

Zoologischer Anzeiger
Deutsche Entomologische Zeitschrift
Coleopterologische Rundschau (Wien)
Annales Zoologici (Warszawa)

Herausgeberschaft

Handbook of Zoology/Handbuch der Zoologie IV Arthropoda: Insecta (N. P. Kristensen & R. G. Beutel)
Handbook of Zoology/Handbuch der Zoologie IV Arthropoda: Insecta, Coleoptera I-IV (R. G. Beutel, R. A. B. Leschen)

Mitgliedschaft in wissenschaftlichen Verbänden:

DZG
Gesellschaft für Biologische Systematik e.V.
Thüringer Entomologenverband
Wiener Coleopterologenverein
Studiengruppe für Zoologische Systematik

             JMorph

Original Articles (Studies published in a Journal without peer review are marked by an asterisk)

Review articles

1. Trautwein, M.D., Wiegmann, B.M., Beutel, R., Kjer, K., & Yeates, D.K. (in press) Advances in Insect Phylogeny at the dawn of the postgenomic era. Annual Review of Entomology.

2. Beutel, R.G. & Schneeberg, K. (in press). Darwin and beetles. Annals of the History and Philosophy of Biology.

3. Beutel, R.G., Leschen, R.A.B. & Friedrich, F. 2009. Darwin, Beetles and Phylogenetics. Naturwissenschaften 96: 1293-1312.

4. Beutel, R.G. & Friedrich, F. 2008. The phylogeny of Archostemata (Coleoptera) and new approaches in insect morphology. Entomologia Generalis 31 (2): 141-154.

5. Beutel, R.G. & Pohl, H. 2006. Endopterygote systematics - where do we stand and what is the goal (Hexapoda, Arthropoda). Systematic Entomology 31: 202-219.

6. Beutel, R.G. 2004. Aktuelle Entwicklungen in der Systematik der Hexapoda (Arthropoda). Entomologie heute 16: 21-50.

 

In press

7. Hünefeld, F. & Beutel, R.G. (in press) The female postabdomen of the enigmatic Nannochoristidae (Insecta: Mecopterida) and its phylogenetic significance. Acta Zoologica.

8. Dallai, R., Mercati, D., Gottardo, M., Machida, R., Mashimo, Y. & Beutel, R. G. (in press) The male reproductive system of Zorotypus caudelli Karny (Zoraptera): sperm structure and spermiogenesis. Arthropod Structure and Development.

9. Jałoszyński, P., Hünefeld, F., Beutel, R.G. (in press) The evolution of "deformed" brains in ant-like stone beetles (Scydmaeninae, Staphylinidae). Arthropod Structure and Development.

10. Dallai, R., Mercati, D., Gottardo, M., Machida, R., Mashimo, Y. & Beutel, R.G. (in press) The fine structure of the female reproductive system of Zorotypus caudelli Karny (Zoraptera). Arthropod Structure and Development.

11. Hünefeld, F., Friedrich, F., Hörnschemeyer, T., Pohl, H. & Beutel, R.G. (in press) The phylogeny of Holometabola and the role of morphology in the age of phylogenomics. DGaaE Nachrichten.

 

2011

12. Schneeberg, K., Courtney, G.W. & Beutel, R.G. 2011. Adult head structures of Deuterophlebiidae (Insecta), a highly derived "ancestral" dipteran lineage. Arthropod Structure and Development 40(1): 93-104. link to pdf

13. Ge, S.-Q., Hörnschemeyer, T., Friedrich, F. & Beutel, R.G. 2011. Is Crowsoniella relicta really a cucujiform beetle? Systematic Entomology 36 (1): 175-179. link to pdf

14. Wipfler, B., Machida, R., Müller, B. & Beutel, R. G. 2011. On the head morphology of Grylloblattodea (Insecta) and the systematic position of the order - with a new nomenclature for the head muscles of Neoptera. Systematic Entomology 36: 241-266. link to pdf

15. Zarinkamar, N., Yang, X., Bao, R., Friedrich, F., Beutel, R., Friedrich, M. 2011. The Pax gene eyegone facilitates repression of eye development in Tribolium. EvoDevo. 2 (1): 8 (15 pp.). link to pdf

16. Friedemann, K. Wipfler, B., Bradler, S. & Beutel, R. G. 2011. On the head morphology of Phyllium and the phylogenetic relationships of Phasmatodea (Insecta). Acta Zoologica, early view DOI: 10.1111/j.1463-6395.2010.00497.x link to pdf

17. Hünefeld, F., Marvaldi, A., Müller, B., Lawrence, J.F. & Beutel, R. G. 2011. The male postabdomen of the "ancestral" archostematan beetle Tetraphalerus bruchi Heller, 1913 (Ommatidae) and its phylogenetic significance. Arthropod Structure and Development 40 (23): 146-158. link to pdf

18. Mashimo, Y., Machida, R., Dallai, R. Gottardo, M. Beutel, R.G. 2011. Egg structure of Zorotypus caudelli Karny (Insecta, Zoraptera, Zorotypidae).  Tissue & Cell 43 (4): 230-237.

19. Ge, S.-Q., Daccordi, M, Beutel, R. G., Li W.-Z &, Yang, X.-K. 2011. Revision of the chrysomeline genera Potaninia, Suinzona and Taipinus (Coleoptera) from eastern Asia, with a biogeographic scenario for the Hengduan mountain region in south-western China. Systematic Entomology 36 (4): 644-671. link to pdf

20. Dressler, C., Ge, S.-Q. & Beutel, R.G. 2011. Is Meru a specialised noterid (Coleoptera, Adephaga)? Systematic Entomology 36 (4): 705-712. link to pdf

21. Hünefeld, F., Pohl, H., Wipfler. B., Beckmann, F. & Beutel, R.G. 2011. The male postabdomen and genital apparatus of †Mengea tertiaria, a strepsipteran amber fossil (Insecta). Journal of Zoological Systematics and Evolutionary Research. early view doi:10.1111/j.1439-0469.2011.00628.x link to pdf

22. Anton, E., Niederegger, S. & Beutel, R. G. 2011. Beetles and flies collected on pig carrion in an experimental setting in Thuringia and their forensic implications. Medical & Veterinary Entomology. early view DOI: 10.1111/j.1365-2915.2011.00975.x link to pdf

 

2010

23. Polilov, A.A. & Beutel, R.G. 2010. Developmental stages of the hooded beetle Sericoderus lateralis (Coleoptera: Corylophidae) with comments on the phylogenetic position and effects of miniaturization. Arthropod Structure and Development 39: 52-69. link to pdf

24. Beutel, R.G., Friedrich, F. & Aspöck, U. 2010. The larval head of Nevrorthidae and the phylogeny of Neuroptera (Insecta). Zoological Journal of the Linnean Society 158: 533-562. link to pdf

25. Friedrich, F. & Beutel, R.G. 2010. The thoracic morphology of Nannochorista (Nannochoristidae) and its implications for the phylogeny of Mecoptera and Antliophora. J. Zool. Syst. Evol. Res. 48: 50-74. link to pdf

26. Schneeberg, K. & Beutel, R.G. 2010. The adult head structures of Tipulomorpha (Diptera, Insecta) and their phylogenetic implications. Acta Zoologica. link to pdf

27. Friedrich, F. & Beutel, R.G. 2010. Good bye Halteria? The thoracic morphology of Endopterygota (Insecta) and its phylogenetic implications. Cladistics 26: 579-612. link to pdf

28. Beutel, R.G, Zimmermann, D., Krauß, M., Randolf, S., & Wipfler, B. 2010. Head morphology of Osmylus fulvicephalus (Osmylidae, Neuroptera) and its phylogenetic implications. Organisms, Diversity and Evolution 10: 311-329.

29. Dressler, C. & Beutel, R.G. 2010. The morphology and evolution of the adult head of Adephaga (Insecta, Coleoptera). Arthropod Systematics and Phylogeny 68: 239-287. pdf

30. Beutel, R. G., Friedrich, F., Hörnschemeyer, T., Pohl, H., Hünefeld, F., Beckmann, F., Meier, R., Misof, B., Whiting, M. F. & Vilhemsen, L. 2010. Morphological and molecular evidence converging upon a robust phylogeny of the megadiverse Holometabola. Cladistics 26: 1-15. link to pdf

31. Ge, S.-Q., Friedrich, F. & Beutel, R.G. 2010. On the systematic position and taxonomic rank of the extinct myxophagan †Haplochelus (Coleoptera). Insect Systematics and Evolution 41 (4): 329-338 link to pdf

32. Pohl, H., Wipfler, B., Grimaldi, D., Beckmann, F. & Beutel, R. G. 2010. Reconstructing the anatomy of the 42 million-year-old fossil †Mengea tertiara (Insecta, Strepsiptera). Naturwissenschaften 97: 855-859. link to pdf


2009

114. Bernhard, D., Ribera, I., Komarek, A. & Beutel, R. G. 2009. Phylogenetic analysis of Hydrophiloidea (Coleoptera: Polyphaga) based on molecular data and morphological characters of adults and immature stages. Insect Systematics & Evolution 40 (1): 3-41. pdf

113. Beutel, R. G., Friedrich, F. & Leschen, R. A. B. 2009. Charles Darwin, Beetles and Phylogenetics. Naturwissenschaften 96 (11): 1293-1312. pdf

112. Beutel, R. G., Kristensen, N. P. & Pohl, H. 2009. Resolving insect phylogeny: The significance of cephalic structures of the Nannomecoptera in understanding endopterygote relationships. Arthropod Structure & Development 38 (5): 427-460. pdf

111. Bravo, F., Pohl, H., Silva-Neto, A. & Beutel, R. G. 2009. Bahiaxenidae, a "living fossil" and a new family of Strepsiptera (Hexapoda) discovered in Brazil. Cladistics 25 (6): 614-623. pdf

110. Eberhard, M. J. B., Pass, G., Picker, M. D., Beutel, R. G., Predel, R. & Gorb, S. N. 2009. Structure and funtion of the arolium of Mantophasmatodea (Insecta). Journal of Morphology 270 (10): 1247-1261. pdf

109. Friedrich, F., Farell, B. D. & Beutel, R. G. 2009. The thoracic morphology of Archostemata and the relationships of the extant suborders of Coleoptera (Hexapoda). Cladistics 25 (1): 1-37. pdf

108. Neugart, C., Schneeberg, K. & Beutel, R. G. 2009. The morphology of the larval head of Tipulidae (Diptera, Insecta) - The dipteran groundplan and evolutionary trends. Zoologischer Anzeiger 248 (3): 213-235. pdf

107. Polilov, A. A. & Beutel, R. G. 2009. Miniaturisation effects in larvae and adults of Mikado sp. (Coleoptera: Ptiliidae), one of the smallest free-living insects. Arthropod Structure & Development 38 (3): 247-270. pdf

106. Xiaoyun, Y., Weber, M., Zarin Kamar, N., Posnien, N., Friedrich, F., Wiegand, B., Beutel, R.G., Damen, W., Bucher, G., Klinger, M. & Friedrich, M. (2009) Probing the Drosophila retinal determination gene network in Tribolium (II): the Pax6 genes eyeless and twin of eyeless. Developmental Biology 333 (1): 215-227. pdf

2008

105. Balke, M., Ribera, I., Beutel, R. G., Viloria, A., Garcia, M. & Vogler, A. P. 2008. Systematic placement of the recently discovered beetle family Meruidae (Coleoptera, Dytiscoidea) based on molecular data. Zoologica Scripta 37 (6): 647-650. pdf

104. Beutel, R. G. & Baum, E. 2008. A longstanding entomological problem finally solved? Head morphology of Nannochorista (Mecoptera, Insecta) and possible phylogenetic implications. Journal of Zoological Systematics and Evolutionary Research 46 (4): 346-367. pdf

103. Beutel, R. G. & Friedrich, F. 2008. The phylogeny of Archostemata (Coleoptera) and new approaches in insect morphology. Entomologia Generalis 31 (2): 141-154.

102. Beutel, R. G. & Friedrich, F. 2008. Comparative study of larval head structures of  Megaloptera (Hexapoda). European Journal of Entomology 105 (5): 917-938. pdf

101*. Beutel, R. G. & Friedrich, F. 2008. A renaissance of insect morphology. DGaaE Nachrichten 22 (1): 5-8. pdf

100. Beutel, R. G., Friedrich, F. & Whiting, M. F. 2008. Head morphology of Caurinus (Boreidae, Mecoptera) and its phylogenetic implications. Arthropod Structure & Development 37 (5): 418-433. pdf

99. Beutel, R. G. & Ge, S.-Q. 2008. The larval head of Raphidia (Raphidioptera, Insecta) and its phylogenetic significance. Zoology 111 (2): 89-113. pdf

98. Beutel, R. G., Ge, S. & Hörnschemeyer, T. 2008. On the head morphology of Tetraphalerus, the phylogeny of Archostemata and the basal branching events in Coleoptera. Cladistics 24 (3): 270-298. pdf

97. Beutel, R. G. & Gorb, S. 2008. Evolutionary scenarios for unusual attachment devices of Phasmatodea and Mantophasmatodea (Insecta). Systematic Entomology 33 (3): 501-510. pdf

96. Beutel, R. G., Krogmann, L., Vilhelmsen, L. T. 2008. The larval head morphology of Xyela sp. (Xyelidae, Hymenoptera) and its phylogenetic implications. Journal of Zoological Systematics and Evolutionary Research 46 (2): 118-132. pdf

95. Beutel, R. G., Ribera, I. & Bininda-Emonds, O. R. P. 2008. A genus-level supertree of Adephaga (Coleoptera). Organisms, Development & Evolution 7 (4): 255-269. pdf

94. Friedrich, F. & Beutel, R. G. 2008. The thorax of Zorotypus (Hexapoda, Zoraptera) and a new nomenclature for the musculature of Neoptera. Arthropod Structure & Development 37 (1): 29-54. pdf

93*. Friedrich, F. & Beutel, R. G. 2008. Micro-computer tomography and a renaissance of insect morphology. Proceedings of SPIE 7048: 1-6.

92*. Friedrich, F., Pohl, H., Hünefeld, F., Beckmann, F., Herzen, J. & Beutel, R. G. 2008. SRμCT-based study of external and internal structures of adults and larvae of Endopterygota (Hexapoda). Hasylab Annual Report 2007: 1527-1528. pdf

91. Pohl, H. & Beutel, R. G. 2008. The evolution of Strepsiptera (Hexapoda). Zoology 111 (4): 318-338. pdf


2007

90. Baum, E., Dressler, C. & Beutel, R. G. 2007. Head structures of Karoophasma sp. (Mantophasmatodea) with phylogenetic implications. Journal of Zoological Systematics and Evolutionary Research 45 (2): 104-119. pdf

89. Baum, E., Hertel, W. & Beutel, R. G. 2007 The head capsule, central nervous system and head circulatory system of an aberrant orthopteran, Prosarthria teretrirostris (Caelifera, Hexapoda). Zoology 110 (2): 147-160. pdf

88. Beutel, R. G., Bocak, L. & Bocakova, M. 2007. Are Polyphaga (Coleoptera) really a basal neopteran lineage - a reply to Kazantsev. Acta Zoologica 88 (2): 153-158. pdf

87. Beutel, R. G. & Vilhelmsen, L. B. 2007. Head anatomy of Xyelidae (Hexapoda: Hymenoptera) and phylogenetic implications. Organisms, Development & Evolution 7 (3): 207-230. pdf

86. Ge, S., Beutel, R. G. & Yang, X.-K. 2007. Thoracic morphology of adults of Derodontidae and Nosodendridae and its phylogenetic implications (Coleoptera), Systematic Entomology 32 (4): 635-667. pdf

85*. v. Knorre, D. & Beutel, R. 2007. Die geowissenschaftlichen Sammlungen im Phyletischen Museum in Jena. Beiträge zur Geologie von Thüringen, NF 13: 61-72.

84. Komarek, A. & Beutel, R. G. 2007. Phylogenetic analysis of Anacaenini (Coleoptera: Hydrophilidae: Hydrophilinae) based on morphological characters of adults. Systematic Entomology 32 (2): 205-226. pdf

83*. Krogmann, L., v. Knorre, D. & Beutel, R. G. 2007. Die Chalcidoidea-Sammlung von Ferdinand Rudow (1840-1920) im Phyletischen Museum (Jena). Entomologische Mitteilungen aus dem Zoologischen und Institut Museum Hamburg 104: 129-140.

 

2006

82. Anton, E. & Beutel, R. G. 2006. On the head morphology of Lepiceridae (Coleoptera: Myxophaga) and the systematic position of the family and suborder. European Journal of Entomology 103 (1): 85-95. pdf

81. Bernhard, D., Schmidt, C., Korte, A., Fritzsch, G. & Beutel, R. G. 2006. From terrestrial to aquatic habitats and back again - molecular insights into the evolution and phylogeny of Hydrophiloidea (Coleoptera) using multigene analyses. Zoologica Scripta 35 (6): 597-306. pdf

80. Beutel, R. G., Balke, M. & Steiner, W. E. 2006. On the systematic position of Meruidae (Coleoptera, Adephaga) and the phylogeny of the smaller hydradephagan families. Cladistics 22 (2): 102-131. pdf

79. Beutel, R. G. & Gorb, S. N. 2006. A revised interpretation of the evolution of attachment structures in Hexapoda (Arthropoda), with special emphasis on Mantophasmatodea. Arthropod Systematics & Phylogeny 64 (1): 3-25. pdf

78. Beutel, R. G. & Pohl, H. 2006a. Head structures of males of Strepsiptera (Hexapoda) with emphasis on basal splitting events within the order. Journal of Morphology 267 (5): 536-554. pdf

77. Beutel, R. G. & Pohl, H. 2006b. Endopterygote systematics - where do we stand and what is the goal (Hexapoda, Arthropoda)? Systematic Entomology 31 (2): 202-219. pdf

76. Friedrich, F. & Beutel, R. G. 2006. The pterothoracic skeletomuscular system of Scirtoidea (Coleoptera: Polyphaga) and its implications for the relationships of the beetle suborders. Journal of Zoological Systematics and Evolutionary Research 44 (4): 290-315. pdf

75. Komarek, A. & Beutel, R. G. 2006. Problems in taxonomy and suggestions for a standardized description of new insect taxa. Entomological Problems 36 (2): 55-70.


2005

74. Balke, M., Ribera, I. & Beutel, R. G. 2005. The systematic position of Aspidytidae, the diversification of Dytiscoidea (Coleoptera, Adephaga) and the phylogenetic signal of third codon positions. Journal of Zoological Systematics and Evolutionary Research 43 (3): 223-242. pdf

73. Beutel, R. G. & Friedrich, F. 2005. Comparative study of larvae of Tenebrionoidea (Cucujiformia, Coleoptera), European Journal of Entomology 102 (2): 241-164. pdf

72. Beutel, R. G. & Leschen, R. A. B. 2005. Phylogenetic analysis of Staphyliniformia (Coleoptera) based on characters of larvae and adults. Systematic Entomology 30 (4): 510-548. pdf

71. Beutel, R. G., Pohl, H. & Hünefeld, F. 2005. Strepsipteran brains and effects of miniaturization. Arthropod Structure & Development 34 (3): 301-313. pdf

70. Beutel, R. G. & Weide, D. 2005. Cephalic anatomy of Zorotypus hubbardi (Hexapoda: Zoraptera): New evidence for a relationship with Acercaria. Zoomorphology 124 (3): 121-136. pdf

69. Hünefeld, F. & Beutel, R. G. 2005. The sperm pumps of Strepsiptera and Antliophora (Hexapoda). Journal of Zoological Systematics and Evolutionary Research 43(4): 297-306. pdf

68. Navarrete-Heredia, J. L., Cortés-Aguilar, J. & Beutel, R. G. 2005. New findings on the enigmatic beetle family Lepiceridae (Coleoptera: Myxophaga). Entomologische Abhandlungen 62 (2): 193-201. pdf

67. Pohl, H. & Beutel, R. G. 2005. The phylogeny of Strepsiptera (Hexapoda). Cladistics 21 (4): 328-374. pdf

66. Pohl, H., Beutel, R. G. & Kinzelbach, R. 2005. Protoxenidae fam. n. (Insecta, Strepsiptera) from Baltic amber - a 'missing link' in strepsipteran phylogeny. Zoologica Scripta 31 (1): 123-134. pdf

 

2004

65. Alarie, Y., Beutel, R. G. & Watts, C. H. S. 2004. Larval morphology of three species of Hygrobiidae (Coleoptera: Adephaga: Dytiscoidea) with phylogenetic considerations. European Journal of Entomology 101 (2): 293-311. pdf

64. Anton, E. & Beutel, R. G. 2004. On the head morphology and systematic position of Helophorus (Coleoptera: Hydrophiloidea: Helophoridae). Zoologischer Anzeiger 242 (4): 313-346. pdf

63. Beutel, R. G. 2004. Aktuelle Entwicklungen in der Systematik der Hexapoda (Arthropoda). Entomologie heute 16: 21-50.

62. Beutel, R. G. & Komarek, A. 2004. Comparative study of thoracic structures of adults of Hydrophiloidea and Histeroidea with phylogenetic implications (Coleoptera, Polyphaga). Organisms Diversity & Evolution 4 (1-2): 1-34. pdf

61. Korte, A., Ribera, I., Beutel, R. G. & Bernhard, D. 2004. Interrelationships of Staphyliniform groups inferred from 18S and 28S rDNA sequences, with special emphasis on Hydrophiloidea (Coleoptera, Staphyliniformia). Journal of Zoological Systematics and Evolutionary Research 42 (4): 281-288. pdf

60. Leschen, R. A. B. & Beutel, R. G. 2004. Ocellar atavism in Coleoptera: plesiomorphy or apomorphy? Journal of Zoological Systematics and Evolutionary Research 42 (1): 63-69. pdf

59. Pohl, H. & Beutel, R. G. 2004. Fine structure of adhesive devices of Strepsiptera (Insecta). Arthropod Structure and Development 33 (1): 31-43. pdf

 

2003

58. Beutel, R. G., Anton, E. & Jäch, M. A. 2003. On the evolution of adult head structures and the phylogeny of Hydraenidae (Coleoptera, Staphyliniformia). Journal of Zoological Systematics and Evolutionary Research 41 (4): 256-275. pdf

57. Beutel, R. G. & Raffaini, G. B. 2003. A new record of Sphaeriusidae for Argentina (Coleoptera: Myxophaga). Coleopterologische Rundschau 73: 1-6.

56. Schunger, I., Beutel, R. G. & Britz, R. 2003. Morphology of immature stages of Prostomis mandibularis (Prostomidae, Tenebrionoidea, Coleoptera). European Journal of Entomology 100 (3): 357-370. pdf

 

2002

55. Beutel, R. G. & Hörnschemeyer, T. 2002a. Larval morphology and phylogenetic position of Micromalthus debilis LeConte (Coleoptera: Micromalthidae). Systematic Entomology 27 (2): 169-190. pdf

54. Beutel, R. G. & Hörnschemeyer, T. 2002b. Description of the larva of Rhipsideigma raffrayi (Coleoptera, Archostemata), with phylogenetic and functional implications. European Journal of Entomology 99 (1): 53-66. pdf

53. Gorb, S., Beutel, R. G, Gorb, E. V., Jiao, Yuekan, Kastner, V., Niedereger, S., Popov, V. L., Scherge, M., Schwarz, U. & Vötsch, W. 2002. Structural design and biomechanics of friction-related releasable attachment devices in insects. Integrative Comparative Biology 42 (6): 1127-1139. pdf

52. Grebennikov, V. V. & Beutel, R. G. 2002. Morphology of the minute larva of Ptinella tenella, with special reference to effects of miniaturisation and the systematic position of Ptiliidae (Coleoptera: Staphylinoidea). Arthropod Structure & Development 31(2): 157-172. pdf

51. Hörnschemeyer, T., Beutel, R. G. & Pasop, F. 2002. Head structures of Priacma serrata Leconte (Coleoptera, Archostemata) inferred from x-ray tomography. Journal of Morphology 252 (3): 298-314. pdf

50. Ribera, I., Beutel, R. G., Balke, M. & Vogler, A. P. 2002. Discovery of Aspidytidae, a new family of aquatic Coleoptera. Proceedings of the Royal Society, London B 269 (1507): 2351-2157. pdf

 

2001

49. Beutel, R. G., Anton, E. & Bernhard, D. 2001. Head structures of adult Spercheus: Their function and possible significance to staphyliniform phylogeny. Annales Zoologici 51 (4): 473-484.

48. Beutel, R. G. & Gorb, S. 2001. Ultrastructure of attachment specializations of hexapods (Arthropoda): evolutionary patterns inferred from a revised ordinal phylogeny. Journal of Zoological Systematics and Evolutionary Research 39 (4): 177-207. pdf

47. Beutel, R. G. & Slípinski, S. A. 2001. Comparative study of larval head structures of Sphindidae and Protocucujidae (Cucujoidea, Coleoptera). European Journal of Entomology 98 (2): 219-232. pdf

46. Gorb, S. & Beutel, R. G. 2001. Evolution of locomotory attachment pads of hexapods. Naturwissenschaften 88 (12): 530-534. pdf

45. Haas, F. & Beutel, R. G. 2001. Wing folding and the functional morphology of the wing base in Coleoptera. Zoology 104 (2): 123-141. pdf

44. Leschen, R. A. B. & Beutel, R. G. 2001. Pseudotracheal tubes, larval head, and mycophagy in Sepedophilus (Coleoptera: Staphylinidae: Tachyporinae). Journal of Zoological Systematics and Evolutionary Research 39 (1-2): 25-35. pdf

 

2000

43. Beutel, R. G. & Haas, F. 2000. Phylogenetic relationships of the suborders of Coleoptera (Insecta). Cladistics 16 (1): 103-141. pdf

42. Beutel, R. G. & Pollock, D. A. 2000. Larval head morphology of Phycosecis litoralis (Pascoe) (Coleoptera, Phycosecidae) with phylogenetic implications. Invertebrate Taxonomy 14 (6): 825-835. pdf

41. Beutel, R. G., Weide, D. & Bernhard, D. 2000. Characters of the larval head of Mycetina cruciata (Schaller) (Coleoptera; Endomychidae) and their phylogenetic implications. Annales Zoologici 50 (1): 7-14.

40. Gorb, S. & Beutel, R. G. 2000. Head-capsule design and mandible control in the beetle larva: A three-dimensional approach. Journal of Morphology 244 (1): 1-14. pdf

39. Jäch, M. A., Beutel, R. Díaz, J. A. & Kodada, J. 2000. Subgeneric classification of Hydraena Kugelann (Coleoptera: Hydraenidae). Annalen des Naturhistorischen Museums Wien 102B: 177-258.

 

1999

38*. Beutel, R. G. 1999a. Description of the larva of Satonius kurosawai (Sato, 1982) (Coleoptera: Torridincolidae). Water Beetles of China 2: 53-59.

37. Beutel, R. G. 1999b. Phylogenetic analysis of Myxophaga (Coleoptera) with a redescription of Lepicerus horni (Lepiceridae). Zoologischer Anzeiger 237: 291-308.

36. Beutel, R. G. 1999c. Morphology and evolution of the larval head structures of Hydrophiloidea and Histeroidea (Coeloptera: Staphylinidae). Tijdschrift voor Entomologie 142: 9-30.

35. Beutel, R. G., Maddison D. R. & Haas, A. 1999. Phylogenetic analysis of Myxophaga (Coleoptera) using larval characters. Systematic Entomology 24 (2): 1-23. pdf

 

1998

34. Beutel, R. G. 1998. Trachypachidae and the phylogeny of Adephaga (Coleoptera). Phylogeny and Classification of Caraboidea. XXI I.C.E. (1996, Firenze). Museo regionale de Scienze Naturali Torino, Atti 1998: 81-106.

33. Beutel, R. G. & Haas, A. 1998. Larval head morphology of Hydroscapha natans LeConte, 1874 (Coleoptera, Myxophaga, Hydroscaphidae) with special reference to miniaturization. Zoomorphology 118 (2): 103-116. pdf

 

1997

32. Beutel, R. G. 1997. Über Phylogenese und Evolution der Coleoptera (Insecta), insbesondere der Adephaga. Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg NF 31: 1-164.

31. Beutel, R. & Molenda, R. 1997. Comparative morphological study of larvae of Staphylinoidea (Coleoptera, Polyphaga) with phylogenetic implications. Zoologischer Anzeiger 236 (1): 37-67.

 

1996

30. Beutel, R. G. 1996. Study of the larva of Nosodendron fasciculare (Olivier 1790) (Coleoptera: Nosodendridae) with implications to the phylogeny of Bostrichiformia. Journal of Zoological Systematics and Evolutionary Research 34 (1): 121-134.

29. Beutel, R. G. & Haas, A. 1996. Phylogenetic analysis of larval and adult characters of Adephaga (Coleoptera) using cladistic computer programs. Entomologica Scandinavica 27 (2): 197-205. pdf

 

1995

28. Arndt, E. & Beutel, R. G. 1995. Larval morphology of Systolosoma Solier and Trachypachus Motschulsky (Coleoptera: Trachypachidae) with phylogenetic consideration. Entomologica Scandinavica 26 (4): 439-446. pdf

27. Beutel, R. G. 1995b. Phylogenetic analysis of Elateriformia (Coleoptera: Polyphaga) based on larval characters. Journal of Zoological Systematics and Evolutionary Research 33 (2): 145-171. pdf

 

1994

26. Beutel, R. G. 1994a. On the systematic position of Hydrotrupes palpalis Sharp (Coleoptera: Dytiscidae). Aquatic Insects 16 (3): 157-164. pdf

25. Beutel, R. G. 1994b. Study on the systematic position of Systolosoma breve Solier (Adephaga: Trachypachidae) based on characters of the thorax. Studies on Neotropical Fauna and Environment 29 (3): 161-167.

24*. Beutel, R. G. 1994c Phylogenetic analysis of Hydrophi­loidea (Coleoptera: Polyphaga: Staphyliniformia) based on characters of the head of adults and larvae. Koleopterologische Rundschau 64, 103-131.

23. Arndt, E. & Beutel, R. G. 1994. Descriptions of the larvae of Paussus (Klugipaussus) aff. distinguendus Reichensberger and P. (Lineatopaussus)afzelii Westwood (Carabidae) with phyloge­netic considerations. Elytron 8: 129-139.

22*. Beutel, R. G. & Jäch, M. 1994. Untersuchungen über die Propleura der Hydraenidae (Coleoptera: Staphyliniformia). Zeitschrift der Arbeitsgemeinschaft Österreichischer Entomologen, 47 Jg. (1/2): 59-63).

 

1993

21. Beutel, R. G. 1993. Phylogenetic analysis of Adephaga (Coleoptera) based on characters of the larval head. Systematic Entomology 18 (2): 127-147. pdf

20. Beutel, R. G. & Roughley, R. E. 1993. Phylogenetic analysis of Gyrinidae based on characters of the larval head (Coleoptera: Adephaga). Entomologica Scandinavica 24 (4): 459-468. pdf

1992

19. Arndt, E., Beutel, R. G. & Joost, W. 1992. Description of the larva of Andogyrus buqueti (Aube, 1838) (Coleoptera, Gyrinidae). Studies of Neotropical Fauna and Environment 28: 139-144.

18. Beutel, R. G. 1992a. Phylogenetic analysis of thoracic structures of Carabidae (Coleoptera). Zeitschrift für zoologische Systema­tik und Evolutionsforschung 30 (1): 53-74. pdf

17. Beutel, R. G. 1992b. On the morphology of the head of the larva of Omoglymmius hamatus (LeConte) and the systematic position of Rhysodidae (Coleoptera, Adephaga). Entomologica Scandinavica 23 (2): 169-184. pdf

16. Beutel, R. G. 1992c. Study on the systematic position of Metriini (Coleoptera, Carabidae) based on characters of the larval head. Systematic Entomology 17 (3): 207-218. pdf

15*. Beutel, R. G. 1992d. External and internal structures of the head of 3rd instar larvae of Carabus coriaceus F. (Adephaga: Carabi­dae). Entomologica Basiliensia 15: 147-168.

14*. Beutel, R. G. 1992e. Larval head structures of Licinus silphoides Rossi (Coleoptera: Carabidae) and their phylogenetic impli­cations. Entomologica Basiliensia 15: 169-194.

 

1991

13. Beutel, R. G. 1991a. Larval head structures of Omophron and their implications for the relationships of Omophronini (Coleoptera: Carabidae). Entomologica Scandinavica 22 (1): 55-67. pdf

12*. Beutel, R. G. 1991b. Internal and external structures of the head of 3 rd instar larvae of Amphizoa lecontei Matthews (Coleoptera: Amphizoidae). A Contribution towards clarification of the systematic position of Amphizoidae. Stuttgarter Beiträge zur Naturkunde (A) 469: 1-24.

 

1990

11. Beutel, R. G. 1990a. Phylogenetic analysis of the family Gyrinidae (Coleoptera) based on meso- and metathoracic characters. Quaestiones entomologicae 26: 163-191.

10. Beutel, R. G. 1990b. Metathoracic features of Omoglymmius hamatus and their significance for classification of Rhysodini (Coleoptera: Adephaga). Entomologica generalis 15 (3): 185-201.

9. Beutel, R. G. & Ruhnau, S. 1990. Phylogenetic analysis of the ge­nera of Haliplidae (Coleoptera) based on characters of adults. Aquatic Insects 12 (1): 1-17. pdf

 

1989

8. Beutel, R. G. 1989a. The head of Spanglerogyrus albiventris Fol­kerts (Coleoptera: Gyrinidae). Contribution towards clarifica­tion of the phylogeny of Gyrinidae and Adephaga. Zoologische Jahrbücher für Anatomie 118: 431-461.

7. Beutel, R. G. 1989b. The prothorax of Spanglerogyrus albiventris Folkerts, 1979 (Coleoptera, Gyrinidae). Contribution towards clarification of the phylogeny of Gyrinidae. Entomologica Basi­liensia 13: 151-173.

 

1988

6. Beutel, R. G. 1988. Studies of the metathorax of the trout-stream beetle, Amphizoa lecontei Matthews (Coleoptera: Amphizoidae): Contribution towards clarification of the systematic position of Amphizoidae. International Journal for Insect Morphology and Embryology 17 (1): 63-81. pdf

5. Beutel, R. G. & Roughley, R. E. 1988. On the systematic position of the family Gyrinidae (Coleoptera: Adephaga). Zeitschrift für zoologische Systematik und Evolutionsforschung 26 (5): 380-400. pdf

 

1987

4. Beutel, R. G. & Roughley, R. E. 1987. On the systematic position of the genus Notomicrus Sharp (Hydradephaga, Coleoptera). Canadian Journal of Zoology 65 (8): 1898-1905.

 

1986

3*. Beutel, R. 1986a. Skelet und Muskulatur des Kopfes und Thorax von Hygrobia tarda (Herbst). Ein Beitrag zur Klärung der phylogene­tischen Beziehungen der Hydradephaga (Insecta: Coleoptera). Stuttgarter Beiträge zur Naturkunde (A) 388: 1-54.

2*. Beutel, R. 1986b. Skelet und Muskulatur des Kopfes der Larve von Haliplus lineatocollis Mrsh. (Coleoptera). Stuttgarter Beiträge zur Naturkunde (A) 390: 1-15.

1*. Beutel, R. & Belkaceme, T. 1986. Comparative studies on the meta­thorax of Hydradephaga and Trachypachidae. Entomologica Basili­ensia 11: 221-229.

 

Books

Beutel, R. G. & Leschen, R. A. B. (eds.) 2005. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim). In: Band/Volume IV Arthropoda: Insecta Teilband / Part 38. Handbook of Zoology. Berlin, New York, De Gruyter.

 

Book contributions

Arce-Pérez, R., Navarete-Heredia, J. L. & Beutel, R. G. 2005. 6.1. Lepiceridae. In: Beutel, R. G. & Leschen, R. A. B. (vol. eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Archangelsky, M., Beutel, R. G. & Komarek, A. 2005. 10. Hydrophiloidea. Introduction, Phylogeny. 10.1. Hydrophilidae. In: Beutel, R. G. & Leschen, R. A. B. (vol. eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Arndt, E., Beutel, R. G. & Will, K. 2005. 7.8. Carabidae. In: Beutel, R. G. & Leschen, R. A. B. (vol. eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Balke M., Ribera, I. & Beutel, R. G. 2003. Aspidytidae: On the discovery of anew beetle family: detailed morphological analysis, description of a second species, and key to fossil and extant adephagan families (Coleoptera). In: Jäch M. A., Ji L. (eds.), Water Beetles of China. Wiener Coleopterologenverein / Vienna Coleopterists Society, Wien, pp. 53-66.

Beutel, R. G. 1995a. The Adephaga (Coleoptera): phyloge­netic analysis and thoughts on the evolutionary history. In: Biology, Phylogeny, and Classification of Coleoptera: Papers celebrating the 80th Birthday of Roy A. Crowson (J. Pakaluk & S. A. Slípinski, eds.). Polish Academy of Science. Warszawa (Muzeum i Instytut Zoologii PAN), pp. 173-217.

Beutel, R. G. 2002. Coeloptera. Archostemata. Adephaga. Myxophaga. In: Lehrbuch der Speziellen Zoologie. Begründet von Alfred Kästner. Band I: Wirbellose Tiere, 5. Teil: Insecta. (JH. Dathe & H.-E. Gruner, eds.). Heidelberg, Berlin (Spektrum Akademischer Verlag GMBH).

Beutel, R. G. 2005. 1. Systematic position and early evolution. 6. Myxophaga. Introduction, Phylogeny. 7.9. Rhysodidae. In: Beutel, R. G. & Leschen, R. A. B. (vol. eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Beutel, R. G. & Arce-Pérez, R. 2005. 6.3. Sphaeriusidae. In: Beutel, R. G. & Leschen, R. A. B. (vol. eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Beutel, R. G. & Arndt, E. 2005. 7.7. Trachypachidae. In: Beutel, R. G. & Leschen, R. A. B. (vol. eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Beutel, R. G. & Lawrence, J. F. 2005. 4. Coleoptera, Morphology. In: Beutel, R. G. & Leschen, R. A. B. (vol. eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Beutel, R. G. & Leschen, R. A. B. 2005. 8. Polyphaga. Introduction, Phylogeny. 9. Staphyliniformia. Introduction, Phylogeny. 18.10. Chelonariidae. In: Beutel, R. G. & Leschen, R. A. B. (vol. eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Beutel, R. G. & Ribera, I. 2005. 7. Adephaga. Introduction, Phylogeny. In: Beutel, R. G. & Leschen, R. A. B. (vol. eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Beutel, R. G. & Roughley, R. E. 2005. 7.1. Gyrinidae. In: Beutel, R. G. & Leschen, R. A. B. (vol. eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Beutel, R. G. & Vanin, S. A. 2005. 6.2. Torridincolidae. In: Beutel, R. G. & Leschen, R. A. B. (vol. eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Costa, C., Vanin, S. A., Ide, S. & Beutel, R. G. 2005. 18.6. Heteroceridae. In: Beutel, R. G. & Leschen, R. A. B. (vol. eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Jäch, M., Beutel, R. G., Delgado, J. A. & Díaz, J. A. 2005. 11.1. Hydraenidae. In: Beutel, R. G. & Leschen, R. A. B. (vol. eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Lee, C.-F., Jäch, M. & Beutel, R. G. 2005. 18.7. Psephenidae. In: Beutel, R. G. & Leschen, R. A. B. (vol. eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Leschen, R. A. B. & Beutel, R. G. 2005. 14. Elateriformia. Introduction, Phylogeny. In Beutel, R. G. & Leschen, R. A. B. (vol. eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

Vanin, S. A., Beutel, R. G. & Arce-Pérez, R. 2005. 6.4. Hydroscaphidae. In: Beutel, R. G. & Leschen, R. A. B. (vol. eds.) Handbook of Zoology, Vol. IV Arthropoda: Insecta. Part 38. Coleoptera, Vol. 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga (partim). Walter De Gruyter, Berlin, New York.

 

Web contents

Tree of life (D. Maddison & W. Maddison): Beutel, R. G. 1996. Haliplidae.

Texte zur Vorlesung 'Einführung in die Zoologische Systematik' (Sommersemester 2008) zum Herunterladen (Word-Dokumente)
(Stand September 2008)

Taxonomie und Systematik
Systematik/Cladismus (Stand 6. Oktober 08)
Geschichte
Fossilien und Erdzeitalter
Evolution und Systematik
Biodiversität (Stand 6. Oktober 08)



Klausur Teilmodul Biodiversität

Termin: Mittwoch, 29.10.08, voraussichtlich 17.15, Großer Hörsaal, Zoologie,  Erbertstraße 1
Möglicherweise ergibt sich aus Raumgründen eine Verzögerung um eine Stunde (18.15)